http://www.timesonline.co.uk/article/0,,60-303790,00.html

Stephen Jay Gould
Evolutionary biologist who challenged the orthodox thinking on Darwinism
and 
had few rivals as a populariser of science
 
 
 
Stephen Jay Gould was one of the most gifted evolutionary scientists of
his 
generation. Following the publication of many eloquently written articles
and 
books, together with numerous public lectures, he acquired a reputation
as an 
outstanding science populariser. In the research field of evolutionary 
biology his reputation was more controversial because of his persistent 
challenging of what he saw as the conventional reductionism of the
orthodoxy, 
with its great emphasis on Darwinian adaptation as the predominant factor
in 
evolution. 

He was born in New York, of second-generation East European Jewish emigre
parents, and took his first degree in geology from Antioch College, Ohio.

Four years of study at Columbia University, involving research on the 
biometrics and evolutionary history of Bermudan Pleistocene land snails,
was 
rewarded with a doctorate, and in 1967 he was appointed assistant
professor 
in invertebrate palaeonotology at Harvard, and assistant curator in the 
Museum of Comparative Zoology. Four years later he was promoted to
associate 
professor, and in 1974 he became a full professor at the unusually early
age 
of 33. 

After moving his principal domicile to New York following his second 
marriage, he took up the post of visiting research professor of biology
at 
New York University in 1996, while maintaining his position at Harvard. 

A longstanding interest in organic growth and form, inspired by the
classic 
work of D'Arcy Thompson, led to the publication of Ontogeny and
Phylogeny, a 
scholarly treatment of the relationship between the growth of individual 
organisms and their evolutionary history. But Gould caused a much greater

stir in evolutionary circles when he and Niles Eldredge propounded the 
hypothesis of punctuated equilibria, which postulates that, contrary to 
conventional Darwinian theory, species exhibit morphological stasis over
long 
periods of time, and give rise to descendent species by means of 
comparatively sudden transformations. 

Just how big a change in evolutionary thought was required to account for

punctuated equilibria has proved debatable but, at the very least, the 
hypothesis directed attention once again to the relevance of the fossil 
record to the study of evolution, at a time when genetics and molecular 
biology were making most of the running. 

In the 1980s Gould went on to promote the idea of species selection to 
account for evolutionary trends recognised in the fossil record, and a 
consequent decoupling of macroevolution (evolution above the species
level) 
from microevolution, as studied by conventional thinkers who, following 
Darwin, accept only selection at the level of the individual. Species 
selection, although theoretically possible, was not well received by 
biologists, and does not receive much empirical support from the fossil 
record; accordingly, it is now generally disregarded. 

With more success Gould challenged other aspects of neo-Darwinism, such
as 
the predominance of adaptive, as opposed to constructional and
historical, 
explanations of organic form. A major theme of his writings in the later 
1980s and 1990s was the key role of historical contingencies in
evolution, 
and the lack of evident progress in general, although he was obliged to 
acknowledge an increase in the complexity of neural systems, culminating
in 
our own species. 

Nevertheless, he considered that human beings might not have evolved but
for 
the chance survival of a primitive chordate ancestor in the Cambrian
period. 
In other words, there was no historic inevitability about our emergence.
This 
is certainly a view that challenges popular wisdom, and it was
popularised in 
his book Wonderful Life. The widespread recognition during this time of
deep 
homologies in the animal world, recognisable at the molecular level,
lends 
support to his belief that internal constraints and channels are
significant 
causes of evolutionary change in their own right, operating to some
extent 
independent of the power of external selection. 

Whatever the dispute that remains about his role as an innovative thinker
in 
evolutionary research, there can be no question about Gould's success as
a 
populariser of science, as recognised by numerous literary awards and 
honorary degrees, to say nothing of a large income derived from this
source, 
which dwarfed his salary as a Harvard professor. In his abundant writings
he 
demonstrated great verbal felicity, a rich vocabulary and capacity for
lucid 
and racy exposition, enlivened by anecdotes, similes and metaphors from 
fields of experience as diverse as baseball and Wagnerian opera. These 
talents were put to effective use for more than a quarter of a century in
a 
series of monthly essays in the magazine Natural History, which concluded

only with the publication of the 300th at the start of what he regarded
as 
the turn of the true millennium, in January 2001. Such was the popularity
of 
these columns that they were anthologised into no fewer than nine books. 

Characteristically, Gould would seize upon some apparently odd feature of

organisms, or quirk of nature, to illustrate, often with great ingenuity,

some evolutionary theme. Some of these essays gave him titles for his
books, 
too, such as The Panda's Thumb or The Flamingo's Smile or Hens' Teeth and

Horse's Toes. Together they show an enviably wide range of learning and 
intellectual curiosity, ranging from homely analogies to the most arcane 
byways of historical scholarship. 

Although predominantly concerned with evolutionary biology, a minority of

them deal with what he saw as the perils of biological determinism.
Always a 
supporter of the underprivileged, Gould was a passionate opponent of 
attempts, conscious or otherwise, by scientists over the past century or
so 
to justify or bolster the entrenched power of the well-educated Caucasian

protestant male in Anglo- Saxon society. He even courted notoriety in the

1970s by allying himself with politically radical groups that were not
always 
scrupulous in their attempts to discredit the newly emergent discipline
of 
sociobiology. 

Gould's social concerns received further expression in The Mismeasure of
Man 
(1981), a tour de force in which he endeavoured to expose the fallacies
and 
concealed biases in a succession of purportedly objective and hence 
influential studies, from mid-19th-century attempts to prove by
craniometry 
the inferiority of North American native peoples and negroes to the
factor 
analytic studies of intelligence by Sir Cyril Burt. Yet Gould never
allowed 
his political radicalism ,” which he espoused sometimes in circumstances
that 
demanded a good measure of personal courage,” to compromise his belief in

individual human rights. Marxism is now long out of fashion but the
beliefs 
he expounded in the prime of his career could perhaps best be described
as 
those of a libertarian Marxist. 

Among the books that made his reputation were Time's Arrow, Time's Cycle 
(1987), a scholarly study of the discovery of geological time, and the 
bestselling Wonderful Life (1989), an account of the remarkable fossil
fauna 
of the Cambrian Burgess Shale in British Columbia, and its evolutionary 
implications. For this he won the Rhône-Poulenc Prize and was
shortlisted for 
the Pulitzer Prize. 

In the following decade his books dealt with topics as varied as
challenging 
the conventional view of evolutionary progress, establishing on good 
scholarly grounds why the new millennium really began in 2001, and
discussing 
the relationship between science and religion. His argument that the two
ways 
of thinking belong to different domains and should be able to co-exist 
without conflict provoked a considerable amount of scepticism, and not
just 
from agnostic or atheistic scientists. 

At nearly 1,500 pages, Gould's most recent book, The Structure of 
Evolutionary Theory, is a summation of his work. It stands by the theory
of 
punctuated equilibria, insisting that it is supported by such fossil
evidence 
as the Burgess Shale, and goes on to reject Richard Dawkins's,selfish
gene 
account of evolution, arguing instead that natural selection occurs on
many 
levels, from the gene to the individual organism, and even the species. 
Finally, it argues against the strict Darwinians that other factors, 
including sheer chance ,” also produce evolutionary change. Reviewing
this 
major contribution to evolutionary theory, in The Times Literary
Supplement 
last week, Steven Rose called Gould the most accomplished living
scientific 
essayist, a match for Haldane in the 1930s and Thomas Huxley in the
latter 
half of the 19th century. 

Among his numerous honours, Gould was one of the first recipients of the 
MacArthur Fellowship (1981-86) and was elected to both the American
Academy 
of Arts and Sciences (1983) and the National Academy of Sciences (1989). 
Perhaps the most distinguished of his many medals was the Gold Medal of
the 
Linnean Society of London, awarded for services to zoology. He even had
an 
asteroid named after him. He served as president of the Palaeontological 
Society in 1985-86, president of the Society for the Study of Evolution 
(1990-91) and president of the American Association for the Advancement
of 
Science (1999-2000). 

A person of strong character and natural ebullience, Steve Gould had
great 
personal warmth and generosity of spirit: unlike some of his radical
allies, 
he was always courteous to his opponents. His interests were
exceptionally 
wide-ranging and his knowledge of many subjects, from medieval
stained-glass 
windows to the history of science, was profound. He had a longstanding 
passionate interest in baseball, and he was able to apply even baseball 
statistics to his intellectual interest in the pursuit of excellence. 

In 1982, when he was gravely ill with asbestos-induced cancer
mesothelioma, 
he was greatly touched to receive a baseball signed by his boyhood hero,
Joe 
DiMaggio. For a short period he even wrote a column on baseball for
Vanity 
Fair. 

He had a good baritone voice and was a keen choral singer. In 1965 he
married 
a fellow Antioch student, Deborah, and after their divorce in 1995 he
married 
Rhonda, a sculptor, and moved to the artists' quarter of Manhattan. He is

survived by his wife and by the two sons of his first marriage. 



Stephen Jay Gould, palaeontologist and popular science writer, was born
on 
September 10, 1941. He died on May 20, 2002, aged 60.
 
 


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