Follow-up Comment #1, bug #319 (project tex4ht):
I've given some information in my answer on TeX.sx:
http://tex.stackexchange.com/a/317390/2891
The problem is that ligatures are formed in this case, hyphens are made active
in \ScriptEnv, the solution is to add \hbox{} to the hyphen definition to
prevent the ligatures forming. I've commited the solution to tex4ht sources.
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