There are two reasons this can happen.

First, and the reason it happens in the examples you linked to, is that in 
most (possibly all) of the built in table of contents macros have layers in 
the hierarchy alternating between bold and normal text. So in that example 
all of the items in the root layer (first, second, third, fourth) are bold 
because they are in the first layer. Then firstone, firsttwo and so on are 
not bold because they are directly below a bolded layer, and then finally 
secondthreeone, secondthreetwo and so on are bold because they are below a 
non-bolded layer.

It is just there to help visually distinguish between the layers in the 
table of contents hierarchy.

Second, if you have plugins that add tiddlers to your table of contents 
those tiddlers will be bold by default because that is how shadow tiddler 
names are displayed. If one of those tiddlers gets changed than the name 
will no longer be bold. This can lead to tiddlers in the table of contents 
not following the above pattern. It is also why I have started moving any 
configuration or changes to tiddlers out of the visible tiddlers in my 
plugins.

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