The parser used in wikitext can be a bit confusing.
There are two important parts here, lists in wikitext are space separated
and things between [[ and ]] are normally interpreted as literal titles
(you show one exception where you have a | that separates the displayed
text from the title it links to).
So [[a title]] being a link to a tiddler and
title [[title with spaces]] somethingElse
is a list of 3 titles.
Both cases are dealing with titles as the atomic units where [[ and ]]
aren't to show that it is a string with spaces, it is to show that it is a
tiddler title. Depending on context a tiddler title can be a link or it can
be an element in a list. Much of what tiddlywiki does is based around lists
and titles.
There are very few limitations on what can be a title, so often anything
between [[ and ]] is considered a literal title. The exception comes from
filters which are actually a superset of the cases where you use [[ and ]]
to denote a title with a string in it.
This explanation could get very long, so to prevent that the answer is that
[[ and ]] is used consistently, but it isn't to show a string, it is to
show a title. In wikitext without a special context like a widget a title
written like that is converted into a link. So your example is being
interpreted as linking to the tiddler with the title {{!!title}} because
otherwise we would have to put restrictions on tiddler naming.
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