Comparing the OSI / IP / Data Property layers.  Upper layers tack on
bytes to the lower layers.

Data=App+Pres+Sess
Segments=Transport
Packets=internet
Frames+Bits=NetworkAccess

MTU is at the layer of frames as in ethernet frames.

MSS is above MTU.  MSS is either at the Packet layer or Segment layer.
 i would think the segment layer.

i am no expert.
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