This is kind of a long post, and many of you will not be interested in it at
all (maybe nobody), but this is in response to the discussion of
antidepressants, depression, its potential causes, and so on. I'm not
turning it in for a grade, so please don't grade me on it. I hope it is
somewhat coherent though, because I really like all of the possibilities for
explanation that emerge in this model. In a previous post, I mentioned
Robert Julien’s *Behavioral Pharmacology* as an excellent resource. Of
course I don’t think he has the final word on the topic of depression and
antidepressants, I do think he provides valuable insight. Paraphrasing what
he has to say on the topic (paraphrasing liberally, I might add, so don’t
fault me for plagiarism—I already know that I am doing that), starting with
the pathophysiology of depression, Julien describes what he refers to as the
neurogenic theory of depression.



This description describes depression as related to loss of volume (due to
neuron loss), especially in the hippocampus.  Julien points out that
numerous symptoms of depression such as impaired attention, concentration,
and memory are related to functions of the hippocampus or the prefrontal
cortex. He also notes that chemicals such as cortisol can have deleterious
effects on the functioning of the hippocampus and prefrontal cortex(tying
together the research that suggests that stress can result in loss of brain
volume, especially in these areas). According to research by Frodl et al.
(2007), the hippocampus actually shrinks when exposed to a variety of
stressors, including depression. That same research reports that stress is
one of the most common causes of depression.



However, Julien notes that recent research has shown that existing neurons
are able to repair or remodel themselves (plasticity), and in fact
neurogenesis also is possible; again, especially in the hippocampus, but
also in the prefrontal cortex. According to his explanation, two known
second messenger neurochemicals  are important in protecting neurons from
damage due to trauma or injury, and in promoting and maintaining the health
and stability of newly generated neurons. Julien argues that antidepressants
increase the action of these 2nd messenger systems resulting in repair and
restoration of volume to the hippocampus and possibly the prefrontal cortex.



Here’s a stepwise summary of how these 2nd messengers might act in the
presence of SSRIs /SNRIs or other antidepressants:



Step 1:  Antidepressant drugs increase the level of the monoamines
(especially 5-HT and NE) in  the synapse.

Step 2: binding of either 5-HT or NE triggers a 2nd messenger system  that
uses  cAMP (cyclicAMP).

Step 3: cAMP triggers the expression of CREB (cAMP response-element-binding
protein).

Step 4: CREB activates the production of BDNF (brain-derived neurotrophic
factor)

Step 5: BDNF promotes healthy neurons and connections (likely involving the
growth of new  neurons, especially in the hippocampus).



Julien supplies ample references to support these statements. Also, Julien
writes that the time frame in which all of these steps occur might explain
the therapeutic lag that accompanies the onset of administration of
antidepressants and the onset of their therapeutic effect.



If you’ve read this far, hopefully I’ve shown the possibility that
depression may be due to stress (loss of a loved one, job loss, ongoing
factors), chemical imbalance (genetic or acquired), or the interaction of
any or all of these factors (the diathesis-stress model). Although
antidepressants may alleviate depression under this model, other things can
as well; for example, psychotherapy, exercise, self-help, stress-reduction
techniques, or time.



Some references:

Angelucci, F., et al. (2005).  BDNF in schizophrenia, depression, and
corresponding animal models. *Molecular Psychiatry* 10, 345-352.

Blendy, J. A. (2006). The role of CREB in depression and Antidepressant
treatments. *Biological Psychiatry, 59,* 1144-1150.



Duman R. S. and Monteggia, L. M. (2006). A neurotrophic model for
stress-related mood disorders. *Biological Psychiatry, 59* , 1116-1127.



Frodl, T. et al. (2007). Association of the brain-derived neurotrophic
factor Val66Met Polymorphism with reduced hippocampal volumes in major
depression. *Archives of General Psychiatry, 64*, 410-416.



Nair, A., and Vaidya, V. A. (2006). CyclicAMP response element binding
protein and brain-derived  neurotrophic factor: Molecules that modulate our
mood? *Journal of Bioscience, 31, *423-434.



Done for the day--going outside as soon as it stops raining.

Carol




-- 
Carol DeVolder, Ph.D.
Professor of Psychology
St. Ambrose University
518 West Locust Street
Davenport, Iowa  52803
563-333-6482

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