Generally, ifupdown is no longer recommended (or supported).  It's
recommended to move to netplan, or at least to directly configuring
networkd or network-manager.

For your specific issue, it's been a while since I looked at ifupdown, but from 
an old bug comment:
https://bugs.launchpad.net/ubuntu/+source/ifupdown/+bug/1628552/comments/9

I think ifupdown static configuration of dns does not work with systemd-
resolved; you must have resolvconf installed for that to work (since
resolvconf installs the ifupdown hook, while resolved ignores ifupdown
completely).  Maybe your system upgrade removed the resolvconf package?


** Changed in: systemd (Ubuntu)
       Status: New => Incomplete

-- 
You received this bug notification because you are a member of Ubuntu
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https://bugs.launchpad.net/bugs/1883412

Title:
  20.04: DNS servers from /etc/network/interfaces not honored by
  systemd-resolve

Status in systemd package in Ubuntu:
  Incomplete

Bug description:
  I have a server that uses /etc/network/interfaces to configure static
  IP info, including DNS, via /etc/network/interfaces. This worked fine
  in 16.04 and 18.04. However, when I updated to 20.04 this no longer
  works; I have to run "systemd-resolve --set-dns=192.168.63.1
  --interface=enp3s0" to fix it after every boot.

  My /etc/network/interfaces is:

  ====================
  # Loopback
  auto lo
  iface lo inet loopback

  # enp3s0: LAN (Motherboard)
  auto enp3s0
  iface enp3s0 inet static
  address 192.168.63.2
  netmask 255.255.255.0
  network 192.168.63.0
  broadcast 192.168.63.255
  gateway 192.168.63.1
  dns-nameservers 192.168.63.1

  iface enp3s0 inet6 static
  address 2001:470:bb52:63::2/64
  netmask 64

  # WiFi USB
  # FIXME: Probably will get renamed on boot
  #allow-hotplug ethFlashAirWifi
  #iface ethFlashAirWifi inet manual
  ====================

  "systemd-resolve --status" outputs the following beforr I run the WAR
  command given above:

  ====================
  Global
         LLMNR setting: no                  
  MulticastDNS setting: no                  
    DNSOverTLS setting: no                  
        DNSSEC setting: no                  
      DNSSEC supported: no                  
            DNSSEC NTA: 10.in-addr.arpa     
                        16.172.in-addr.arpa 
                        168.192.in-addr.arpa
                        17.172.in-addr.arpa 
                        18.172.in-addr.arpa 
                        19.172.in-addr.arpa 
                        20.172.in-addr.arpa 
                        21.172.in-addr.arpa 
                        22.172.in-addr.arpa 
                        23.172.in-addr.arpa 
                        24.172.in-addr.arpa 
                        25.172.in-addr.arpa 
                        26.172.in-addr.arpa 
                        27.172.in-addr.arpa 
                        28.172.in-addr.arpa 
                        29.172.in-addr.arpa 
                        30.172.in-addr.arpa 
                        31.172.in-addr.arpa 
                        corp                
                        d.f.ip6.arpa        
                        home                
                        internal            
                        intranet            
                        lan                 
                        local               
                        private             
                        test                

  Link 4 (tun0)
        Current Scopes: none
  DefaultRoute setting: no  
         LLMNR setting: yes 
  MulticastDNS setting: no  
    DNSOverTLS setting: no  
        DNSSEC setting: no  
      DNSSEC supported: no  

  Link 3 (lxcbr0)
        Current Scopes: none
  DefaultRoute setting: no  
         LLMNR setting: yes 
  MulticastDNS setting: no  
    DNSOverTLS setting: no  
        DNSSEC setting: no  
      DNSSEC supported: no  

  Link 2 (enp3s0)
        Current Scopes: none
  DefaultRoute setting: no  
         LLMNR setting: yes 
  MulticastDNS setting: no  
    DNSOverTLS setting: no  
        DNSSEC setting: no  
      DNSSEC supported: no  
  ====================

  ProblemType: Bug
  DistroRelease: Ubuntu 20.04
  Package: systemd 245.4-4ubuntu3.1
  ProcVersionSignature: Ubuntu 5.4.0-37.41-generic 5.4.41
  Uname: Linux 5.4.0-37-generic x86_64
  ApportVersion: 2.20.11-0ubuntu27.2
  Architecture: amd64
  CasperMD5CheckResult: skip
  Date: Sun Jun 14 01:58:10 2020
  InstallationDate: Installed on 2016-04-12 (1524 days ago)
  InstallationMedia: Ubuntu-Server 16.04 LTS "Xenial Xerus" - Beta amd64 
(20160408)
  MachineType: To Be Filled By O.E.M. To Be Filled By O.E.M.
  ProcEnviron:
   TERM=xterm-256color
   PATH=(custom, no user)
   XDG_RUNTIME_DIR=<set>
   LANG=en_US.UTF-8
   SHELL=/bin/bash
  ProcKernelCmdLine: BOOT_IMAGE=/vmlinuz-5.4.0-37-generic 
root=/dev/mapper/severn_vg2016-lv_root ro
  SourcePackage: systemd
  UpgradeStatus: Upgraded to focal on 2020-06-14 (0 days ago)
  dmi.bios.date: 06/18/2010
  dmi.bios.vendor: American Megatrends Inc.
  dmi.bios.version: P2.40
  dmi.board.name: 890GX Extreme3
  dmi.board.vendor: ASRock
  dmi.chassis.asset.tag: To Be Filled By O.E.M.
  dmi.chassis.type: 3
  dmi.chassis.vendor: To Be Filled By O.E.M.
  dmi.chassis.version: To Be Filled By O.E.M.
  dmi.modalias: 
dmi:bvnAmericanMegatrendsInc.:bvrP2.40:bd06/18/2010:svnToBeFilledByO.E.M.:pnToBeFilledByO.E.M.:pvrToBeFilledByO.E.M.:rvnASRock:rn890GXExtreme3:rvr:cvnToBeFilledByO.E.M.:ct3:cvrToBeFilledByO.E.M.:
  dmi.product.family: To Be Filled By O.E.M.
  dmi.product.name: To Be Filled By O.E.M.
  dmi.product.sku: To Be Filled By O.E.M.
  dmi.product.version: To Be Filled By O.E.M.
  dmi.sys.vendor: To Be Filled By O.E.M.

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