I believe Riseup uses Roundcube as webmail. Last time I checked, both the backend PHP and the client JavaScript code of Roundcube are free software. If Riseup does any bug fixing or feature implementation, I hope they're doing it upstream or they publish their modified version of Roundcube.

I've had a quick look at ProtonMail and it seems to have free JavaScript client code. I believe we can get a copy of that Expat-licensed code by running:

git clone https://github.com/ProtonMail/WebClient.git

I'm not sure if they've released the backend software. Probably not.

(Also, ProtonMail seems to want to provide IMAP/SMTP for desktop clients, but haven't managed yet:
https://protonmail.com/support/knowledge-base/imap-smtp-and-pop3-setup/)

I haven't checked OpenMailBox yet.

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