The bug still lives in 2.28.1.
I found that it should be related with filtering somehow.

In my mailbox settings I turned off 'Apply filters to new messages in Inbox on 
this server', deleted the folders as described above, and after the next 
synchronization I can see all of the missing messages.
I have no local mail filters set up, but when this box was checked, I can see 
my messages while the filtering process was running, but they disappeared 
immediately when this process finished, but still counted as unread in Inbox.

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missing message in the inbox folder
https://bugs.launchpad.net/bugs/160142
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