According to
http://kb.mozillazine.org/Actions_for_attachment_file_types#Unexpected_handling_of_certain_attachments
the problem might arise from a corruption of the mimetypes.rdf file in
the thunderbird profile. In my case simply deleting this file and hoping
that thunderbird will restore it didn't work, though.

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Some attchments (.pdf, .xlsx) don't have open dialog just save dialog
https://bugs.launchpad.net/bugs/593459
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