My system is an upgraded Ubuntu 10.04 (from 09.04).

I had the same error behaviour: cups did not start with boot time
(allthough inserted in /etc/rc.local) but did start manually when
entering /etc/init.d/cups start in the terminal

I found the follwoing solution:
when downgrading the package "ifupdown" from Version 0.7~alpha (which was 
installed during the upgrade) to Version 0.6.8ubuntu29, cups now starts 
automatically again.

-- 
upstart not starting init-scripts (event net-device-up IFACE=lo missing)
https://bugs.launchpad.net/bugs/497299
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