I'm not sure what is happening in your case, but it's working for me in
both Thunderbird and Firefox. Could you please try with a x86_64 build
from ftp://ftp.mozilla.org/pub/thunderbird/nightly/latest-comm-central/
instead of your own build?

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https://bugs.launchpad.net/bugs/732572

Title:
  New Mail Notification Sound does not play in Natty

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