It appears that this is a Sylpheed bug.
The developer advises:
"bug has been fixed at svn r3049.
2012-04-18
* addr_compl.c: fixed Bug #52: Address autocomplete:
Not possible to select drop-down list element with arrow keys
(Linux) Possibly occured with GTK+ >= 2.24.
Patch:
http://sylpheed.sraoss.jp/redmine/projects/sylpheed/repository/revisions/3049/diff/src/addr_compl.c
It was fixed right after the release of 3.2.0beta7,
but it is not released yet.
Please apply the above patch if you are in a hurry."
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https://bugs.launchpad.net/bugs/886593
Title:
Keyboard selection of address autocomplete in email compose window
fails
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