Using Lubuntu 12.04 (and as i recall, same experince with Ubuntu Studio
10.04) -- i have esound AND pulseaudio.  Neither the default new mail
sound nor a custom .wav file would work either in the preferences "test"
dialog nor in actual use until i removed pulseaudio.  I didn't
experience any problem having it in a subfolder of "home."

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https://bugs.launchpad.net/bugs/732572

Title:
  New Mail Notification Sound does not play in Natty

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