I also have the same issue on a large number of PCs. 
The workaround mentioned (@svadim - thank you!) does work. 
It appears that the /tmp/mozilla_mozillaUser0 dir is owned by the first person 
to use it with only owner write permissions set & doesn't get removed at logoff 
or recreated as /tmp/mozilla_mozillaUser# for each additional user.
Setting global write permissions therefore allow any user access to the dir & 
downloads can function correctly.

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https://bugs.launchpad.net/bugs/1527884

Title:
  Firefox 43.0: Downloads fail

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