I believe the answer is because the of ZFS filesystem, including /boot.
Grub can not read ZFS filesystems, my understanding that is in
development.

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https://bugs.launchpad.net/bugs/1873008

Title:
  package lubuntu-grub-theme (not installed) failed to install/upgrade:
  error creating symbolic link './boot/grub/themes/lubuntu-grub-
  theme/icons/ubuntu.png': Operation not permitted - symbolic link on
  vfat filesystem

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