I have no idea whether some of the old, old comments in this bug report
are still relevant or not, but there was no mailto problem between
Firefox and Thunderbird in Fedora 29, or any earlier Fedora release in
recent years that I can recall.  I first saw this symptom in Fedora 31,
which currently has Firefox 75.0 (64-bit) and Thunderbird 68.7 (64-bit).
It does not occur when Chrome is the browser and Thunderbird the default
email client.

It is unclear whether the recent visibility of this symlptom is caused
by underlying changes in Fedora 31, Thunderbird, or Firefox, but since
the error only occurs when both Firefox and Thunderbird are involved,
that points the finger at one or both of those applications.  From the
users viewpoint, these are two independent applications.   Whatever
status information or internal support that Firefox and Thunderbird are
sharing that causes this problem, they obviously shouldn't be sharing.
It should be irrelevant to Firefox's external behavior whether
Thunderbird is the default email client, and irrelevant to Thunderbird
external behavior whether it receives a mailto request from Firefox or
some other browser.

This defect is a major annoyance, as I frequently use web sites with
mailto links, and Firefox and Thunderbird are my preferred applications
for browsing and email.

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https://bugs.launchpad.net/bugs/711305

Title:
  Thunderbird is already running, but is not responding. To open a new
  window, you must first close the existing Thunderbird process, or
  restart your system. Error received when clicking a mailto link in
  firefox.

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