Alright, so this "works for me" [tm] to fix this problem:

Open firefox and enter "about:config" into the location bar.
Set the filter to "mailto". Locate the "network.protocol-handler.app.mailto" 
line, right-click and select "reset".
Close firefox.
Open nautilus/thunar/whatever and go to your firefox profile directory 
(/home/<user>/.mozilla/firefox/<some weird string>/)
There's a file called "mimeTypes.rdf". Remove it (or rename it, if you're the 
cautious type).
Start firefox again.
Right-click anywhere and select "Send Link...".  A window should pop up that 
prompts you to choose your mailer application. Locate /usr/bin/thunderbird (or 
whatever is your preferred app).  Click ok and the compose window should show 
up.

For the curious:
If you now take a look at the about:config page, you won't find the 
network.protocol-handler.app.mailto property anymore. It seems it is not used 
anymore in firefox 3, however having it still set after upgrading seems to 
seriously confuse firefox 3.

-- 
mailto links in Firefox 3.0b5 do not open up Thunderbird compose windows
https://bugs.launchpad.net/bugs/222944
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