Philippe Verdy :

>>Clearly there does seem to be missing a Greek letter,

I hope there is no suggestion of encoding the huge variety of Greek 
abbreviations and ligatures. The early printed Greek texts used type designed 
to follow the manuscripts, but thank goodness that was dropped before long. 
There is a bewildering number of these signs.
See, for example
http://commons.wikimedia.org/wiki/File:Greek_alphabet_ligatures.jpg
and that's just the start of it.

Raymond Mercier

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