oVirt definitely involves a lot of paddling. I see now that you are right about 
HCI (Hyper Converged Platform), it was never specifically documented how to 
upgrade it. I have stepped away from HCI a long time ago, after testing and 
working with it in oVirt 3.x. There were just too many dependencies and things 
that could go wrong. Too much functionality depending on the same hardware and 
software. I have also stepped away from Gluster since and in production we are 
not using self-hosted engines either. A standalone engine on a different 
hypervisor cluster gives me much more peace of mind. However, I cannot see from 
the original post if this is Hyperconverged or even if it is self-hosted. Maybe 
I'm just missing it :-)

The procedures for self-hosted and standalone engine are described and I have 
been able to upgrade clusters in the past using them. Make sure you follow all 
the steps one by one in the correct order. But I agree, if something goes wrong 
on the way, you are a bit on your own in dangerous territory paddling up the 
creek :-P It sounds like the upgrade went fine though... and migrating VMs from 
a 4.3 host to a 4.4 host should be a standard procedure when upgrading the 
nodes one by one in a live environment. The upgrade from 4.3 to 4.4 I cannot 
remember if we did live, but from 4.4 to 4.5 worked for us without taking the 
VMs down. This kind of upgrade is best done in a service window though, and you 
should have backups of everything before you start. I'm 100% with Thomas on 
this.
Looking at "12.6. Migrating hosts and virtual machines from oVirt 4.3 to 4.4" 
from the upgrade guide, I see lots of caveats. It is really depending whether 
or not the oVirt node appliance is used or a Linux Enterprise server (as it 
needs to be upgraded first by the looks of it). And whether or not there are 
VMs with CPU-passthrough.

I have also looked around for bugs in 4.4, as that is the version you are 
upgrading to... I find this for example: Bug 1774064. It seem this error occurs 
in various versions of 4.4, even none related to upgrades.

I'm wondering, if you should not just push through with updating all the oVirt 
nodes, which then allows you to change the cluster compatibility level to 4.4. 
This would mean not migrating VMs live, but shutting them down and possibly 
starting them on an upgraded node, until all nodes are upgraded. If you upgrade 
one more node, you would be able to check if migration works between upgraded 
4.4 nodes, which would then confirm that pushing through and upgrade all nodes 
to 4.4 would be the way forward, even if it means that you have to shutdown all 
the VMs at some point during the migration.

I hope this helps. Good luck with it.

Ps. I have resorted in few cases, to simply reinstalling a node from scratch on 
the new version, and then simply add i to the (upgraded) engine, to make them 
work again. oVirt is a lot of paddling, but in my experience it works fine when 
you just let it run and don't do anything too fancy to it (like upgrading, 
which has to be done from time to time).
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