I know you don't think that this is a DNS problem.  I suggest that you
go over your DNS with a fine tooth comb and if you don't see anything
wrong with it... Get a second opinion.

We had this same problem and it turned out to be the reverse DNS lookup
entries that were incorrect.  The IP numbers were wrong.  They were set
for the old IP address range for the private network so the numbers were
real close and we just plain ol' missed it.

Troy E Bouchard
Network Support

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