OK, I found the answer to my questions :

All is done by Main module :

serve_controller(request, response, session)

source code
this function is used to generate a dynamic page. It first runs all
models, then runs the function in the controller, and then tries to
render the output using a view/template. this function must run from
the [application] folder. A typical example would be the call to the
url /[application]/[controller]/[function] that would result in a call
to [function]() in applications/[application]/[controller].py rendered
by applications/[application]/views/[controller]/[function].html

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