Hi Popko,

My sense is that if the polytomy is hard, it is never necessary to subtract a degree of freedom. If the polytomy is "firm" (sensu Purvis & Garland 1993) - in that the true tree is fully bifurcating but we cannot resolve a multifurcating node due primarily to rapid divergence of the descendant taxa - then subtracting a degree of freedom will probably be excessively conservative. If the polytomy is truly soft (sensu Maddison 1989; Garland & Diaz-Uriarte 1999), then subtracting a degree of freedom for each extra "furc" will probably also be conservative. [As Rohlf 2001 points out, this would imply zero degrees of freedom if the phylogeny was completely unresolved (p. 2154), which seems excessive.] Some of these issues are discussed in Garland & Diaz-Uriarte (1999; Syst. Biol.) and elsewhere.

I'm sure that other subscribers to the list could add significant insight to this issue beyond what I am able to offer.

- Liam

--
Liam J. Revell
University of Massachusetts Boston
web: http://www.bio.umb.edu/facstaff/faculty_Revell.html
(new) email: liam.rev...@umb.edu
(new) blog: http://phytools.blogspot.com

On 2/22/2011 5:31 AM, Popko Wiersma wrote:
Dear all,

Can somebody tell whether one should subtract degrees of freedom when
applying PGLS with a phylogeny containing soft polytomies? And does it
make a difference if polytomies originate from data from multiple
specimens of species?

cheers, Popko Wiersma

- I reposted this message because the first attempt did not show the
text directly in the message body. Apologies for any confusion -

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