Take a m4a file that plays fine on your iPod (I have a old 5th
generation) and change the genre of it in Rhythmbox. After coping the
file back to your ipod (I use gtkpod) your ipod will attempt to play the
song but after a couple seconds it will advance to the next song, never
outputting any sound. For me it displays the track title/artist/album &
album art correctly on the iPod, it appears it is trying to read the
song and just gives up.

This entry from the rhythmbox dev mailling list may be of help:
http://www.nabble.com/Writing-m4a-tags-td20284610.html
"I've committed it, but anyone wanting to try it out should be aware
that m4a files edited in rhythmbox won't play in itunes, and I assume
won't play on ipods either.  I have no idea if this is a problem in
qtmux or something I'm doing wrong, but maybe someone who has more than
ten m4a files will spend the time to figure it out.  I'm not
particularly worried about this as the only people who have qtmux at the
moment are those going out of their way to test it.  Nothing has changed
for those who don't have qtmux installed. "

I opened up both m4a files in a hex editor and the broken one seems to
not only have its metadata changed, but the header is changed as well.

** Attachment added: "After modifying the genre in Rhythmbox"
   http://launchpadlibrarian.net/31106410/broken.m4a

-- 
Editing m4a metadata corrupts file 
https://bugs.launchpad.net/bugs/420893
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