Actually, if you read through the comments of #500417 the scenario when
nautilus doesn't honor the theme is just one special scenario of that
defect and in the comments it is expanded that randomly either the whole
desktop or just the nautilus doesn't honor the theme.

I am on Dell Precision 5500 (dual xeon, 4 core each, no ssd, dual nvidia
quatro with proprietary driver) on 10.10 and I am experiencing the same
problem as you have described "entire gnome desktop reverts to the
default theme on boot, except the theme selector window". The #68
doesn't fixes it and #70 fixes it except the icons on the desktop in
nautilus and which remains still from the old theme.

>From the user perspective the behavior for these two defects is exactly
the same. You have narrowed it down to one hardware configuration where
it always occurs and one workaround which for this hardware
configuration always fixes it, but in my opinion/experience the problem
affects more than just this one hardware configuration and since it is
most likely a race condition or timing issue of some sort, the success
rate of the workaround is just a pure luck and coincidence rather than a
fix.

The underlying issue for both of these defects is exactly same.
Something happens between gnome-systems-settings and nautilus which
causes the entire desktop to revert to the default theme. Changing
timing of the start or restarting gnome-settings-daemon seems to fix
some effects of this problem but not necessarily all of them.

Even if these defects are not duplicates they are at least related since
the underlying issue for both of them is the same.

-- 
Meerkat (10.10) does not apply theme properly
https://bugs.launchpad.net/bugs/649809
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