I have the problem too. I tested about permissions, but it's not the
problem. The problem is with Firefox. I tested with other browser, such
Epiphany, and it was ok.

The problem: cannot save when i put the ftp address directly in the
browser, such http://www.lidec.futuro.usp.br/envio/animagimp.pdf

Cannot save as well when I try to save files from e-mails, for example
Gmail.

The problem does not occur when I firstly open the file with other program, 
such adobe reader, and then save the file. 
 
Also, the problem does not occur when I save in my home directory. The problem 
is for FAT particion. 

I hope this way you have enough information.

-- 
Saving of files only possible in Home folder, not on USB disk, other partitions
https://bugs.launchpad.net/bugs/91348
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