Hi Eric,
 
If your Phaser results show a high Z-score (> 8) AND high LLG AND your solution 
packs without clashes AND refines (even though starting R/Rfree is high) AND 
reproduces density for the model portion AND produces some Fo-Fc density for 
the missing portion, most probably your solution is correct. 
Starting R/Rfree can be high, especially given that you have only ~50% of the 
molecule and which may have structural differences from the target.
 
What you could try out is to manually position the second domain/portion based 
on the partial density and any other information you may have, draw a mask 
around this complete molecule and use this mask for further density 
modification to see if you can improve the density in the missing region. You 
can try several different molecular masks with different positions of the 
missing portion. 
 
Unfortunately, if that does not help, you will have to resort to experimental 
phasing. The phasing power from the partial model may not be good enough to get 
complete phase information for the whole molecule. This should be easier in 
your case assuming the MR solution is right. Even if you get weak/poor 
derivatives, you may be able to find heavy atoms sites using the partial MR 
phases and then combine MR+experimental phases. 
 
Regards,
Debanu.

________________________________

From: CCP4 bulletin board [mailto:[EMAIL PROTECTED] On Behalf Of Eric Liu
Sent: Tuesday, July 31, 2007 2:24 PM
To: [email protected]
Subject: [ccp4bb] how to bring back the missing density for half of the 
structure


Hi All, 
I would like to get some help from here for a data set I recently worked on. I 
have been working on a new kinase data set which does not have a close homolog. 
The data was collected to 2.1A  resolution in space group P212121 however the 
difference between a and b is only 0.5A. If I index the data as P4, Rmerge is 
increased from 13% to 39%. I used the most close homologs which have about 37% 
sequence identity as search model for molecular replacement and it seemed I 
have got the solution by using Phaser with only the c-terminal part of the 
search model and also a long loop removed. After changed the different residues 
back to the target protein, the structure was refined to Rfree/R  46% and 43%  
to 2.1 A resolution. The existing c-terminal structure has well defined density 
except 25ish residue at the very c-terminal end doesn't have well connected 
density. Current model contains about 50% of overall target residues. I can see 
some extented difference density for several residues going
to the N-terminal part and also extented density for the C-terminal loop for 
several residues. I also see tones of not well-conncted difference density in 
the N-terminal region. There was no sever clashes between molecules after mount 
all symmetry related molecules. My question is the following: 
 
1. Have I got the correct  solution for the molecular replacement?
2. How can I bring back the missing density for the N-terminal residues and the 
loop region?
 
I would really appreciate any inputs or suggestions.
 
Eric

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