Dear CCP4bb,

I am attempting to crystallise a 25kDa membrane protein of eukaryotic origin.  
We have obtained crystals of the protein (with and without a potential ligand). 
 However crystal quality is poor, as exposure at room temperature and 
cryo-protected conditions have given diffraction as far as 10-15Ang at best.  
After scouring the CCP4bb archives and local expertise, we have not yet had 
much success in improving crystal quality, and wished to probe the knowledge of 
the community for additional ideas.
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The background;

The protein is purified from its native source, solubilised with 
octyl-glucoside detergent, and treated with a recombinant PNGaseF to remove 
glycosylation.  After an initial Q-sepharose and size exclusion chromatography, 
the protein is concentrated by elution from a small volume of Q-sepharose resin 
(centrifugal concentration is occasionally used, but introduces an unknown in 
the detergent concentration, the monomers of which move through the 
concentrator membrane, but not micelles).

The protein has been reproducibly crystallised in glycine and bicine buffers, 
at low pH ( 9.0-10.0 ), low molarity salt, and 30-33% PEGs of various molecular 
weights (e.g. PEG 300, 1000, 2000).  Native crystals had a size and morphology 
very similar to crystals of a close homologue, appearing sharp-edged and quite 
stable to careful manipulation with cryo-loops.

Crystals obtained in the presence of the potential ligand had a different, less 
sharp morphology (more like thin plates).  Notably the latter crystals seemed 
to form only in what appeared to be a phase separation, but on manipulation 
seemed more like a gel, perhaps protein precipitate.  The gel made the crystals 
difficult to manipulate, and possibly resulted in mechanical damage.  Of more 
concern I believe it may have been looped up with the crystal and prevented 
proper cryoprotectant penetration, although there were no ice rings to indicate 
so.

Neither form diffracted beyond 10Ang, even in different cryoprotectants (higher 
PEG concentration, 25% glycerol, sucrose, and ethylene glycol).  Furthermore 
automated annealing (for 1sec) did not improve diffraction.  Seeding has been 
trialled for the native crystals (not yet for ligand-bound forms), but has not 
improved crystal growth.

The purification detergent used is being reconsidered.  We have previously 
attempted to crystallise the protein in nonyl-glucoside detergent, without 
success.  Various additive detergents (below their CMC), alcohols and other 
amphilic additives have been screened, without success in crystallisation.  We 
aim to swap the protein into different detergents (i.e. maltosides) and try for 
improved crystal quality under known conditions.

We are also considering crystal dehydration, in an attempt to reduce solvent 
content.  Additionally I have attempted reproducing conditions with 0.1% w/v 
agarose as an additive, aiming to promote growth of the latter crystal form 
without the difficult gel phase.  Finally I have toyed a bit with cubic lipid 
phase crystallisation, without any success so far.  Any advice on these 
specific approaches would be most appreciated.

As you can see we have considered many methods.  If there is something I have 
missed, or perhaps some common pitfall I have not anticipated, I would 
appreciate any advice you have to offer.


I thank you for taking the time to read this mini-essay, and again for 
answering my off-topic request for advice.

Regards,
Damon.

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Damon Colbert
School of Biological Sciences
University of Auckland

Email: [email protected]




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