Dear Wei,

The enzyme ribonucleotide reductase can, depending on organism and class, bind 
ATP as a substrate in the active site (c), as an allosteric regulator of 
substrate specificity at another site (s) and as an overall activity regulator 
at a third site (a)! It can also bind dATP at the second and third sites, but 
not as a substrate. It cannot bind to sites (c) and (s) at the same time 
although it could potentially bind to sites (s) and (a) simultaneously. Here is 
a review that you might find useful:

http://www.ncbi.nlm.nih.gov/pubmed/22050358

Best wishes
Derek

On 4 Mar 2014, at 04:48, Wei Shi <[email protected]> wrote:

> Dear all,
> Does anyone happen to know examples of 2 ligands bind to a single protein / 
> each monomer protein in 2 different ligand binding pockets? 
> I know the following example:
> (1). phosphofructokinase, which binds ATP as both a ligand and a feedback 
> inhibitor in different sites 
> (2).  2 cAMP bound to each E. coli CAP monomer in the crystal structure. 
> 
> Does any of you know other examples? Thank you so much!
> 
> Best,
> Wei 

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