There is a really interesting paper, just available online, that speaks 
directly to this question. The paper is about something else but the model 
parameters can be translated into expected life-spans of homonin taxa.

Bokma, F., V. van der Brink, and T. Stadler. In press. Unexpectedly many 
extinct hominins. Systematic Biology. (available online as an "early view" 
paper).

http://onlinelibrary.wiley.com/doi/10.1111/j.1558-5646.2012.01660.x/abstract

-lh


On Jul 19, 2012, at 9:45 AM, Neahga Leonard wrote:

> Hi William,
> 
> As far as I understand it, most of the studies on past organisms are done
> at the genus level, not the species level, making accurate evaluations of
> species-life-spans exceedingly difficult.  The fuzziness of where the line
> is drawn dividing where one species becomes another adds yet another level
> of difficulty.
> 
> The fossil record bias toward preserving remains of larger animals rather
> than smaller ones would also seem to impart a potentially heavy bias in
> estimating species-life-spans as well.
> 
> Within the homo/Australopithecus lineage, *H. erectus*, has had the longest
> run so far, with most of the other branches of the family being relatively
> short-lived.  I don't know of any agreement as to why this is.  I've heard
> suggested that it's a product of poor fossil record, that it is a
> reflection of a period of rapid evolution with many different forms
> speciating rapidly, then dying off, and that it is an artifact of
> artificially creating different species categories.
> 
> The article below indicates a mammal average species-lifespan within the
> time-frame you've mentioned, but that estimate may be subject to some of
> the biases I mentioned earlier.
> 
> "Mammals are among the fastest-radiating groups, being characterized by a
> mean species lifespan of the order of 2.5 million years (Myr)1,
> <http://www.nature.com/nature/journal/v443/n7112/full/nature05163.html#B1>2<http://www.nature.com/nature/journal/v443/n7112/full/nature05163.html#B2>.
> The basis for this characteristic timescale of origination, extinction and
> turnover is not well understood."
> 
> http://www.nature.com/nature/journal/v443/n7112/abs/nature05163.html
> 
> Good luck with your search, I'd be very curious to learn what you discover.
> 
> Neahga Leonard
> 
> On Thu, Jul 19, 2012 at 10:37 AM, William Grove-Fanning <
> [email protected]> wrote:
> 
>> To clarify: I
>> mean the the lifespan of the sub-species Homo sapiens sapiens, not the
>> life-span of its
>> individual members.  I believe that for species of larger mammals, it is
>> around 1-2
>> million years, but need to firm up and be able to cite any such number.
>> 
>> Thanks Again, William
>> 
>>> Date: Wed, 18 Jul 2012 10:16:39 -0500
>>> From: [email protected]
>>> Subject: [ECOLOG-L] Lifespan of the Modern Human (Homo Sapiens Sapiens)
>>> To: [email protected]
>>> 
>>> I am doing research for a paper that requires a fairly general  answer
>> to what appears to be, at least on the face of it, a fairly simple
>> question.  But I am having a heck of a time finding any sort of answer.
>> Here is the question:
>>> 
>>> Has anyone estimated or tried to estimate the life span  of the modern
>> human (homo sapiens sapiens)?
>>> 
>>> The question, of course, gets muddied quickly with the possibility of
>> genetic manipulation and other self-directed evolutionary interventions; the
>>> creation of isolated human populations in space; and a global disaster
>> that wipes out all life on earth.  But assuming a “base rate” of human (or
>> mammalian or large mammalian) speciation, how long is the modern human, as
>> a sub-species, expected to be around?  Alternatively, what is the average
>> life span of members of the genus Homo (e.g., Homo sapiens, Neanderthals,
>> Homo erectus, etc.)?
>>> 
>>> 
>>> 
>>> If someone could point me to a citable source, I would reallyappreciate
>> it.  Thanks!  William
>>> 
>>> - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -
>> - - - - -
>>> William Grove-Fanning, Andrew W. Mellon Fellow
>>> Department of Philosophy, Environmental Studies Program, Trinity
>> University
>>> Secretary-elect, Newsletter Editor, & Website Manager, International
>> Society for Environmental Ethics
>>> 
>>> 
>> 

Luke Harmon
Assistant Professor
Biological Sciences
University of Idaho
208-885-0346
[email protected]

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