Wikipedia actually has a fairly good resource on this:
http://en.wikipedia.org/wiki/List_of_animals_displaying_homosexual_behavior

I haven't looked into the sources in detail, and I'm sure the definition is
nuanced in each case, but it's a good starting point and it's fully
referenced.

As for maintainence and ultimate explanations of homosexuality in nature...
A more interesting hypothesis than the 'inclusive fitness' argument (in my
opinion) is that genes coding for homosexuality are maintained through
inheritance in female lines: those which manifest in males as homosexual
orientation are manifest in female relatives to increase fitness. There's
some (relatively anecdotal) evidence supporting this:
http://www.medicaldaily.com/articles/10287/20120613/homosexuality-gene-mother-reproduction-evolution.htm

Iain

- - - - - - - - - - - -
Dr. Iain Stott
Environment and Sustainability Institute
University of Exeter, Cornwall Campus
Tremough, Treliever Road
Penryn, Cornwall, TR10 9EZ, UK.
- - - - - - - - - - - -
http://www.exeter.ac.uk/esi/
http://biosciences.exeter.ac.uk/cec/



On 28 March 2013 03:40, Warren W. Aney <[email protected]> wrote:

> Wayne and Ecolog-L,
> I recall reading some time ago about a hypothesis that homosexuality in
> populations is or may be a response to heightened population density,
> implying that it is an innate stratagem to reduce reproduction while
> maintaining adult productivity.  Anyone know anything about this?
>
> Warren W. Aney
> Tigard, Oregon
> (503) 539-1009
>
>
> -----Original Message-----
> From: Wayne Tyson [mailto:[email protected]]
> Sent: Wednesday, 27 March, 2013 11:53
> To: Warren W. Aney; [email protected]
> Subject: Re: [ECOLOG-L] Expedition notice and question
>
> Warren and Ecolog:
>
> As a stock-farmer's son, I have seen plenty of what appears to us to be
> homosexual behavior among, for example, cows and bulls and ducks and dogs,
> but I have never witnessed actual penetration, nor have I witnessed any
> cases where bulls eschew cows in favor of sex with bulls. Homosexual in the
> sense that it occurs in humans is the center of my interest, and anything
> that is relevant to that question is of interest to me, including
> anecdotes.
>
> "Mounting" of other bulls has long been considered dominance behavior, and
> this occurs in humans who consider themselves heterosexual, as appears to
> be
>
> the case with dogs.
>
> Your point is well made about the continuum; Alan Watts once wrote an
> article entitled (as I recall) "The Circle of Sex." That, or anything else
> I
>
> have read, does not address my intentionally restricted question.
>
> WT
>
> ----- Original Message -----
> From: "Warren W. Aney" <[email protected]>
> To: "'Wayne Tyson'" <[email protected]>; <[email protected]>
> Sent: Wednesday, March 27, 2013 10:30 AM
> Subject: RE: [ECOLOG-L] Expedition notice and question
>
>
> > Wayne, I once had a beef cow that refused to mate with a bull -- she did
> > lactate and help nurture another cow's calf.  The cow may have been
> > homosexual or asexual.  I've heard livestock owners say that cows
> > frequently
> > display homosexual behavior (mounting other cows) but a cow exhibiting
> > excessive homosexual behavior including avoiding bills is usually sold
> for
> > slaughter.
> > As I understand it, in nature (including humans) there is a wide and
> > continuous spectrum of sexual behavior ranging from pure heterosexuality
> > to
> > bisexuality to pure homosexuality, and this range of behaviors is further
> > modified by a varying continuum of sexual intensity from hypersexuality
> to
> > asexuality.
> >
> > Warren W. Aney
> > Tigard, Oregon
> > (503) 539-1009
> >
> >
> > -----Original Message-----
> > From: Ecological Society of America: grants, jobs, news
> > [mailto:[email protected]] On Behalf Of Wayne Tyson
> > Sent: Wednesday, 27 March, 2013 10:32
> > To: [email protected]
> > Subject: [ECOLOG-L] Expedition notice and question
> >
> > [NOTE:] I will be on expedition (with a stop at the National Native Seed
> > Conference in Santa Fe NM on April 10) until the two weeks at the end of
> > April and the first week of May, then gone again beginning the 2nd week
> of
> > May until around May 24. I will not be checking email during those
> > periods,
> > but will respond to as many email messages as possible during those
> > hiatuses. A third expedition following those is likely, but the period of
> > hiatus is iffy.]
> >
> > Here is my parting question. Please feel free to post it on other lists.
> >
> > Re: Homosexuality in animals other than Homo sapiens. We know that
> > homosexual behavior occurs in other species in some forms (Bonobo
> > chimpanzees [Pan paniscus], for example), and we know that hermaphrodites
> > of
> > some species fertilize each other simultaneously. But my question is in
> > which species other than humans, does EXCLUSIVE homosexuality, especially
> > in
> > the form of pair bonds, occur?
> >
> > WT
> >
> > I'll pick up my answers in late April. If I have time, I may be able to
> > respond to some today. Please respond on-list, and not to me personally.
> >
> >
> >
> > -----
> > No virus found in this message.
> > Checked by AVG - www.avg.com
> > Version: 10.0.1430 / Virus Database: 2641/5707 - Release Date: 03/27/13
> >
>

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