I'm (well) over my head and (well) outside whatever wheelhouse I might occupy but I am compelled:

I think part of why this keeps circling is that we’re staying at the level of language—what counts as an “it,” whether something qualifies as an “invasive species.” Not unimportant, but  downstream of the actual dynamics?

What shows up first, at least in the field examples glen mentioned, isn’t a species, it’s a change:

 * crops failing
 * pollination dropping
 * something spreading

A shift in the landscape—some affordance opening up, some constraint disappearing.

“invasive species” reads less like a fundamental category and more like a label we apply post-hoc.  Based on an identified a carrier that matters to us. Anthrorelativism creeps in—we tend to call something “invasive” when the change is fast /and/ intersects with human interests?

I’m more interested in the mismatch in timescales:

 * ecosystems can reconfigure niches relatively quickly (disturbance,
   climate, land use, etc.)
 * species/genetic adaptation is significantly slower

you get situations where the niche landscape shifts faster than the resident community can track, and then whatever competency bundle happens to fit—whether already present or newly arrived—expands rapidly.

At that point, asking “what is the invasive species?” feels a bit like asking for the name of the thing after the phase transition has already started. Sometimes there is a clear “it,” sometimes there isn’t, or it’s a constellation.

So I’m less concerned with pinning down the ontology up front, and more with:

 * what changed in the affordance structure
 * how quickly it changed
 * and which populations were able to respond on that timescale

The “invasive” label then becomes a kind of shorthand for that high-rate, human-salient reconfiguration, rather than the starting point of the explanation.

It also is deeply loaded with judgement, reflecting the "convenience" to the observer?

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