PS, David, I'm aware of the 500-million-year decline you point to, but
that's not what I was talking about.  Each warming has had active
biological cycling.

Evolution has worked quickly compared to the pace of the ice ages;
each ice age cycle has had newly evolved organisms involved in the
cycling of carbon

Take the coccolithophores as one major example; sorry I don't know
where to find a copy but this cite gets at how they changed from
coastal to pelagic organisms, taking over the open ocean and so having
far more ability to cycle carbon:

de Vargas, C.; Aubry, M.P.; Probert, I.; Young, J.  (2007). Origin and
evolution of coccolithophores: from coastal hunters to oceanic
farmers, in: Falkowski, P.G.; Knoll, A.H. (Ed.) (2007). Evolution of
primary producers in the sea. pp. 251-285

This example is an old favorite article of mine:
http://courses.washington.edu/ocean450/Discussion_Topics_Papers/Schmitz_et_al.pdf
(as corrected:  
http://www.nature.com/nature/journal/v407/n6803/full/407467a0.html
)


"... certain species of plankton characteristic of high-productivity
regions flourished all over the world at the P/E boundary6. .... a
large peak in biogenic barite ... at the P/E boundary indicates a
dramatic increase in biological productivity7. Biogenic barite has
proved a reliable proxy for surface-water biological productivity in
the open oceans of the past7,8.
The paper by Bains et al.1 not only provides new evidence that oceanic
productivity did indeed increase, but also provides a feasible
mechanism for how an episode of greenhouse warming may end. .... Bains
et al. propose that higher productivity and the resulting sequestering
of excess carbon in the oceans, through photosynthesis, was the
feedback mechanism required to bring levels of atmospheric CO2 and
temperatures back to normal...."

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