One of the things we found when photographing in European museums is that
the rules are very strict on who the copyright of the photographed image
belongs to. It definitely isn't the photographer. In some places where there
is a strict no-photography rule (Verulamium, Wallace Collection etc), we
were able to photograph providing that we signed a form saying that we would
only used the photographs for research purposes, not for commercial use. Any
copyright strictly stayed with the museum.

A publisher is held legally responsible for clearing permissions to copyrighted works used in the publication. The photo copyright could belong to a photographer (one who did not sign the relevant rights away to a museum), or to a museum, or to someone hired by the publisher or author. However, the publisher always knows who the copyright belongs to, and the author often knows. If the publisher does not hold the copyright they can direct you to the relevant party, and the author can always direct you to the publisher.

Fran

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