I have read the preceding arguments with some interest.  I have played 
principal, second, and fourth horn professionally (occasionally subbing on 
third, plus an extra like 5th, 7th or even Wagner tuba) in orchestras and 
played horn 1-4 in professional bands.  If this were a band, I would probably 
recommend third, but for orchestra, things are more complicated.  One thing, 
however, that should not sway your decision is the prevalence of solos.  In an 
orchestra, all of the horn parts are solo parts and require solo capable 
players (in other words, in a modern orchestra everyone must be able to play 
everything in the entire range of the instrument).  As I have written before, 
the literature has many solos for all horn parts.  Using second horn as an 
example, look through the Beethoven symphonies, remember that the Rite of 
Spring starts with solo bassoon and second horn, Ein Heldenleben's opening horn 
solo is restated by the second hornist 16 measures later, down a minor third!  
 Here is the way that most of the hornists in orchestras that I have played in 
view the standard horn section:
 
Principal (also sometimes called Solo horn): Provides style, intonation and 
section leadership.
Second horn:  Shares brain stem with Principal.  If the principal provides the 
style, the second horn either makes or breaks it (another way of saying this 
is: The principal provides the style; the second horn sets the style).  Playing 
second is a lot like playing catch* with the principal horn.
Third horn:  Either is a second principal horn (if the music is paired with 
horns 1&2 in one key and 3&4 in another), assistant to the principal (many 
times the principal will lay out if the third doubles first), or provides the 
loudness in loud high playing.  Of course, sometimes the third player is just 
the third player (Richard Deane wrote a great article for The Horn Call about 
this).
Fourth horn:  Either shares brain stem with the 3rd horn or provides the 
foundation of the quartet sound.
 
I am aware that some people may find this controversial.  This is, however, 
what works in the region that I live and play in.  As an additional disclaimer, 
I won the fourth horn in the orchestra I play in. When the second horn position 
opened up, I auditioned and won the second position (which I currently hold).  
When the third horn position opened up, I did not audition, even though it may 
have increased my chance to sub as principal for one simple reason:  We play 
more stuff for only two horns then there are opportunities to sub as principal. 
 In other words, the pay (and play!) is better.
 
Respectfully Submitted,
Scott Young
 
* "Catch" is an American game of tossing a (base)ball back and forth.           
                          
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