Cf. Coleman ad loc. (/e/ = short e; /E/ = long e) "tul/e/runt: but tul/E/runt at 5.34. -/e/runt, the older form, is more frequent in iambic and other metres, but rare in hexameters owing to the preponderance of perfect stems ending in long vowels; cf. miscueruntque (G. 3.283), constiterunt (A. 3.681)."
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