Dear colleagues,

On behalf of my co-authors, I am pleased to announce the publication of the 
following papers on subantarctic fur seal fasting capabilities and their 
ecological implications. If you are interested in PDF copies, please email me 
at: [email protected]

 

Best wishes,

 

Delphine

 

 

PAPER #1:

 

Verrier, D., Groscolas, R., Guinet, C. and Arnould, J. P. Y. , Development of 
fasting abilities in subantarctic fur seal pups: balancing the demands of 
growth under extreme nutritional restrictions. Functional Ecology, no. doi: 
10.1111/j.1365-2435.2010.01823.x

 

Summary:

 

1. Surviving prolonged food deprivation requires various metabolic adaptations 
such as energy and protein sparing, which can be highly conflicting with 
energy-demanding stages of an animal’s life history such as growth.

2.Due to the maternal attendance pattern, subantarctic fur seal (Arctocephalus 
tropicalis Gray) pups must repeatedly endure exceptionally long fasts of 
increasing duration throughout the 10-month lactation period. Little is known 
of (i) how these infants adapt to such extreme energetic constraints while 
sustaining growth and development; and (ii) the ecological implications of 
repeated prolonged fasting in early life in terms of offspring survival, 
maternal care and growth strategy in this species, as well as the evolutionary 
consequences of such life history trait.

3. Physiological responses  to prolonged fasting and how they change with 
development throughout the pre-weaning period were investigated. Results show 
that beginning with their first fast, subantarctic fur seal pups are able to 
mobilize lipid reserves preferentially while conserving protein stores in 
response to nutritional deprivation. As pup age, profound changes in energy 
expenditure allow the implementation of an efficient strategy of fat storage 
and lean body mass preservation, which proves highly adaptive in the face of 
the low maternal provisioning rates experienced.

4. Despite increasing fasting durations, pup mortality decreased markedly 
throughout the maternal dependence period. Consistent with predictions, field 
measurements indicate that fasting endurance, although limited in early life, 
increases up to durations of nearly 3 months with age. Results suggest that the 
maternal provisioning strategy could be constrained by these ontogenetic 
changes in pup fasting abilities.

5. Furthermore, extreme energetic constraints and local density-dependent 
effects appear to exert a strong selective pressure upon the adoption of a 
convergent growth strategy between the sexes aiming to maximize fat storage and 
pre-weaning survival.

6. The issues of resulting trade-offs between pre- and post-weaning survival 
and the evolutionary consequences of extreme fasting abilities are also 
addressed.

 

 

PAPER #2:

 

Verrier, D., Guinet, C., Authier, M., Tremblay, Y., Schaffer, S., Costa, D. P., 
Groscolas, R. and Arnould, J. P. , The ontogeny of diving abilities in 
subantarctic fur seal pups: developmental trade-off in response to extreme 
fasting?. Functional Ecology, no.doi: 10.1111/j.1365-2435.2011.01846.x

 

Summary:

 

1. A major hypothesis of life-history theory is that conditions of early 
development affect future survival and reproductive success. Responses to 
detrimental environments during early ontogeny may involve trade-offs between 
current and future fitness. Yet, the functional mechanisms involved in such 
evolutionary trade-offs remain poorly documented.

2. The physiological and behavioural ontogeny of diving abilities was examined 
in subantarctic fur seal (Arctocephalus tropicalis Gray) pups to assess whether 
the repeated extreme fasts they naturally endure throughout the period of 
maternal dependence impacts on their development.

3. The ontogeny of pup body oxygen storage capacity was slow, in particular for 
the muscle compartment, which shows limited increase in myoglobin content 
(0·23–0·85 g 100 g wet muscle−1). As a consequence, by the time of weaning, 
mass-specific oxygen stores had only reached 76%, 24% and 61% of adult female 
capacity for blood, muscle and total, respectively. Concomitantly, in marked 
contrast to other pinniped infants, they spent decreasing amounts of time in 
water (16–7%) with age and exhibited very little diving experience and skills.

4. Overall, in addition to experiencing the longest fasting durations 
throughout the maternal dependence period, subantarctic fur seal pups 
demonstrate the lowest levels of mass-specific total oxygen stores and maximum 
dive duration of any otariid near the age of weaning reported to date.

5. Furthermore, dives that exceeded the calculated aerobic dive limit occurred 
with a very low frequency (0·04%), suggesting that behavioural limitations 
linked to the necessity to conserve energy to survive repeated fasting, rather 
than restricted oxygen storage capacity, constrained pup diving behaviour.

6. Hence, these animals appear to trade-off the early development of both their 
physiological and behavioural diving abilities in favour of body fat 
accumulation to survive the prolonged fasts they must endure and, potentially, 
provide a nutritional buffer while they locate appropriate food patches after 
weaning. While promoting pre-weaning survival, this strategy renders pups more 
vulnerable to unpredictable changes in environmental conditions and food 
availability at the transition to independent foraging and, thus, could have 
negative impact on post-weaning survival.

 

*********************************************************************************

Dr Delphine Verrier

DVM, PhD

Head of Primate Centre

Projet leader for the "Primatology and Health" programme (URES)

International Centre for Medical Research of Franceville (CIRMF)

B.P. 769, Franceville, Gabon

Phone : +241 67 70 62 / Mobile: +241 07 98 56 69

Email:  <mailto:[email protected]> [email protected]

Skype: ddlafouine1

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cid:34994B7F-E15C-441B-8A1E-AFDCC209897F

 

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