Dear colleagues,

My co-authors and I are excited to announce the publication of our new
paper, “Diving Behavior of Common Bottlenose Dolphins Around the Main
Hawaiian Islands: Individual Variation and Responses to Diel and Lunar
Cycles”, in Marine Mammal Science.
Please feel free to request a PDF copy from us via email at
*[email protected]* <[email protected]>. You can
find the abstract (in both English and ʻŌlelo Hawaiʻi) below, and at
*https://onlinelibrary.wiley.com/doi/10.1111/mms.70232*
<https://onlinelibrary.wiley.com/doi/10.1111/mms.70232>. You can also
download a copy of the supplemental materials at
*https://cascadiaresearch.org/files/Harnish_etal_2026MMS_supplemental.pdf*
<https://cascadiaresearch.org/files/Harnish_etal_2026MMS_supplemental.pdf>


Annette Harnish

Research Biologist

Cascadia Research Collective

*www.cascadiaresearch.org* <http://www.cascadiaresearch.org>

CITATION:
Harnish, A.E., R.W. Baird, and M.A. Kratofil. 2026. Diving Behavior of
Common Bottlenose Dolphins Around the Main Hawaiian Islands: Individual
Variation and Responses to Diel and Lunar Cycles. Marine Mammal Science
42(3): e70232. doi: 10.1111/mms.70232

ABSTRACT:
Marine mammals navigate their environment both horizontally and vertically,
adding an additional layer of complexity to understanding their behavior
and developing effective conservation strategies. Here, we use data from 14
tagged common bottlenose dolphins (*Tursiops truncatus*) from Hawai’i to
explore how their diving behavior varies in response to temporal and
demographic variables. Overall mean dive depth (for dives ≥50m) was 206m
(max=896m), and duration was 5.9min (max=13.6min). However, individual
variation was considerable and explained most of the variation in dive
depths and durations. Dive rates varied between diel periods, with rates at
night more than double those during the day. Dive depths and durations also
varied between diel periods, with animals generally conducting their
deepest and longest dives during dusk and their shortest and shallowest
dives at night. Regular dives to or close to the seafloor occurred in
depths of up to 800m, but animals spent the majority of their time in the
top 50m of the water column. These findings represent the first baseline
data on diving behavior for this species in Hawai’i and will be critical in
evaluating potential impacts stemming from planned marine infrastructure
development.

HŌ‘ULU‘ULU MANA‘O:
Hoʻokele nā māmala kai i ko lākou kaiapuni ma ka papakū me ka papamoe, a he
kūʻonoʻono ke ʻano o ka maopopo ʻana i kā lākou lawena, i ko lākou ʻano, a
pehea e hoʻolālā a hoʻomōhala aku ai i mau kaʻakālai maluō kūpono. I kēia
pepa, kālailai aku mākou i ka ʻikepili mai ʻumikūmāhā mau naiʻa nuku
poko (*Tursiops
truncatus*) o Hawaiʻi i lepili ʻia i mea e kālailai aku ai i ke ʻano o ko
lākou luʻu ʻana a pehea e loli ai i hāpane i nā kumuloli kaime a henua. ʻO
ka ʻawelike hohonu o ka luʻu ʻana (no nā luʻu i ʻoi aʻe a like paha me ke
50m), he 206m (ka hohonu loa=896m), a ʻo ka lōʻihi he 5.9 minuke (ka lōʻihi
loa=13.6 minuke). He nui nō naʻe nā ʻaui ʻana ma nā pākahi, a ʻo ia nō ke
kumu nui o nā ʻaui ʻana ma ka hohonu a me ka lōʻihi o ka luʻu ʻana. Ua pā
ka heluna o ka luʻu ʻana i ka lā, ʻoi aku, ma ka pālua hoʻi, ka nui o ka
heluna luʻu ma ka pō ma mua o ke ao. Pēlā pū ka hohonu a me ka lōʻihi o nā
luʻu ʻana, aia nō i ka māhele o ka lā, a ma ka maʻamau, helu ʻia nā luʻuna
hohonu a lōʻihi loa ma ka mōlehulehu, a ʻo nā luʻuna pōkole a pāpaʻu loa ma
ka pō. Helu ʻia nā luʻuna maʻamau i kokoke i ka papakū ma ka hohonu a hiki
i ka 800m, noho naʻe ma kahi o nā 50m o luna o ke kolamu kai i ka hapanui o
ka manawa. ʻO kēia mau hua nā ʻikepili kahua mua ma ke ʻano o ka luʻu ʻana
no kēia lāhui ma Hawaiʻi, a he mau ʻikepili waiwai a koʻikoʻi nō ma ka
loiloi ʻana i nā hopena e pā aku ana paha kēia ʻano kaiāulu i ke kūkulu
kumuhana kai e hoʻomākaukau ʻia ana.
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