Damien

I should also say that given the range of size variation you are suggesting
(i.e. where there is no overlap in size distributions between the focal
species for instance ) then I would not use the approach I suggested. You
would be in a position of extrapolating the shape ~ size relationship
likely too far away from where you would be comfortable with the prediction
(and uncertainty in the prediction).

Also if your intraspecific shape ~ size models are for ontogenetic
allometries I am even more skittish (because ontogenetic allometries so
often have non-linear relationships). So it may be that what I advocated is
not useful for your questions.

On Tue, 14 Jul 2020 at 21:32, Damien Esquerre <[email protected]>
wrote:

> Dear all,
> thanks for the excellent feedback. I need to maybe clarify some things:
>
> I am definitely aware and agree that interspecies comparisons need to be
> done in a phylogenetic context. However, when you have multiple individuals
> per species, and are comparing intraspecific vectors such as allometric
> slopes between species (I know it sounds contradictory but hopefully you
> know what I mean) I am not aware of any method that can incorporate
> phylogenetic information (these usually rely on species means, or one value
> per species). We sort of explored a bit of that issue in the attached
> paper. I guess I can think of ways around it, like including 'clade' as a
> factor in the model, but that doesn't fully account for the relationships
> between species. I think Dean and Mike have been working on this?
>
> I like Ian's suggestion of using species means at a comparable size.
> However, that wouldn't work when we are talking about species with orders
> of magnitude in size difference.
>
> Something I can think of, even if ontogenetic trajectories are different
> between species, one could compute the evolutionary allometric trajectory
> of the mean adult shape of a group of species, and then extract the
> residuals from that regression. I guess there could be many arguments
> against doing this if those species have different ontogenetic
> trajectories, but would love to open a discussion about this.
>
> Another option is just not performing any allometric corrections and
> accept there will be a confounded allometric component to variation with
> the evolutionary (interspecific variation).
>
> In the end, what would you do if you wanted to detect mode of evolution,
> convergence and or adaptation (effect of environment) in a clade that
> displays heterogeneous allometric slopes? In particular, when you dont have
> comprehensive ontogenetic series and that is not the focus of your question.
>
> This is something I've been thinking about for years, and it fascinates
> me, but have never arrived at a satisfactory answer.
>
> Thanks again!
> Damien
>
> On Wed, Jul 15, 2020 at 6:04 AM Mauro Cavalcanti <[email protected]>
> wrote:
>
>> Dear Hugo,
>>
>> >shape of one particular species. Now regarding the Ian example in
>> Drosophila, I tested allometry into 60 species across the genus and there
>> is definitely a >pattern (indeed a beautiful one across the genus looking
>> into the different clades) but looking into one particular group of species
>> the "size" is indeed a very
>>
>> Have you already published this work? If so, could you please provide a
>> reference? It looks truly great.
>>
>> With best wishes,
>>
>> Em ter., 14 de jul. de 2020 às 16:57, Hugo Benítez <
>> [email protected]> escreveu:
>>
>>> Fantastic line of discussion
>>>
>>> I'm absolutely agree with Joe, when there is not a Phylogenetic context
>>> maybe the allometric correction would not have very much sense, because we
>>> are looking into only one generation so we really don't know very well if
>>> the shape we are looking for is product of environmental condition (that
>>> can be connected with plenty of variables like nutrition, stress, etc...)
>>> now as you asked there is multiple group of species so definitely in your
>>> idea there is some "historic factor" that provide the shape of one
>>> particular species. Now regarding the Ian example in Drosophila, I tested
>>> allometry into 60 species across the genus and there is definitely a
>>> pattern (indeed a beautiful one across the genus looking into the different
>>> clades) but looking into one particular group of species the "size" is
>>> indeed a very good trait to explain differences (like species from island,
>>> marsh or the typical cosmopolitan) cosmopolitan Drosophila have the simple
>>> small wing (very small). On the other hand, where maybe "makes sense"  is
>>> in one single species after doing some quantitative genetics experiments
>>> and controlling the factors that could influence the size depending on your
>>> question...   But I think if there is a simple species in the game the
>>> factors in one generation are indeed just the real biological meaning of
>>> your differences and I dont think a correction will have a very biological
>>> meaning...   Now of course could be some exception to the rule and a
>>> Biogeographical question like bergmann rule, or another rule like that
>>> where the relationship is directly related to size maybe a correction could
>>> be ok to see how big there are the differences when the factor is
>>> included...
>>>
>>> I would love to see more replies,  nice topic to discuss Damien
>>>
>>> Best
>>> Hugo Benítez
>>>
>>>
>>> El mar., 14 jul. 2020 a las 14:55, Damien Esquerre (<
>>> [email protected]>) escribió:
>>>
>>>> Dear morpho community,
>>>> I have a philosophical question on size correction that should start an
>>>> interesting discussion.
>>>> When we are interested in seeing the effects of species or
>>>> environmental variables for example, on shape, people often first remove
>>>> allometric variation by computing the residuals of a shape ` size
>>>> regression. This of course, doesn't make sense if there are heterogeneous
>>>> slopes and species have different allometric trajectories (i.e. if the
>>>> species*size term is significant).
>>>> What do you think would be the most appropriate way to deal with this
>>>> situation then, if you are interested in environmental effects on shape?
>>>> Best regards,
>>>> Damien Esquerré
>>>>
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>>>> .
>>>>
>>>
>>>
>>> --
>>>
>>> *Dr. Hugo A. Benítez*
>>> Profesor Asociado
>>> Centro de Investigación de Estudios Avanzados del Maule
>>> Universidad Católica del Maule
>>>
>>> Research Associate, University of Cambridge Museum of Zoology
>>> External Researcher Faculty of Agriculture, University of Zagreb
>>>
>>> Lab website: http://www.morphoshape.com <http://www.hugoabenitez.com>
>>>
>>>
>>>
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>>> .
>>>
>>
>>
>> --
>> Dr. Mauro J. Cavalcanti
>> E-mail: [email protected]
>> Web: http://sites.google.com/site/maurobio
>> "Life is complex. It consists of real and imaginary parts."
>>
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>


-- 
Ian Dworkin
Department of Biology
McMaster University
Office phone 905 525 9140 ext. 21775
Lab phone 905 525 9140 ext. 20076
[email protected]
dworkinlab.github.io

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