Hi Pablo

As an alternative to considering just half the skull, you can use the whole 
skull and extract the symmetric component of shape variation with an 
analysis of object symmetry. This was published by Kanti Mardia and 
colleagues in to somewhat technical papers:

Mardia, K. V., F. L. Bookstein, and I. J. Moreton. 2000. Statistical 
assessment of bilateral symmetry of shapes. Biometrika 87:285300.
Kent, J. T., and K. V. Mardia. 2001. Shape, Procrustes tangent projections 
and bilateral symmetry. Biometrika 88:469485.

There is also a paper introducing this approach in a form that should be 
more accessible to biologists (I hope):

Klingenberg, C. P., M. Barluenga, and A. Meyer. 2002. Shape analysis of 
symmetric structures: quantifying variation among individuals and 
asymmetry. Evolution 56:19091920.
(PDF file posted on my web site, http://www.flywings.org.uk )

The approach adds a reflected and relableled copy of the original 
configurations to the originals, and then includes both in a Procrustes 
analysis. You can do the reflection and relabeling steps in any statistics 
package that allows data manipulation or in a spreadsheet. The Procrustes 
analysis can then be done with any of the usual morphometrics programs.

This approach avoids the problem with the landmarks on the median plane 
that you have with the analysis of half-skulls. Also, unlike other methods 
of reflecting and averaging the paired landmarks, you don't have to choose 
some median landmarks of which you assume that they are exactly in the 
median plane.

I hope this helps.

Best wishes,
Chris



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Christian Peter Klingenberg
Faculty of Life Sciences
The University of Manchester
Michael Smith Building
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United Kingdom

Telephone: +44 161 275 3899
E-mail: [EMAIL PROTECTED]
Web: http://www.flywings.org.uk
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