application = ErrorMiddleware(application, debug=True)

That line is sure to cause problems. Debugging Middleware usually
causes issues with mod_wsgi because unless you run a very specific
configuration to eliminate multi-threaded requests, though, the error
you're getting is that application isn't properly being passed to your
ErrorMiddleware.

I think you also need to remove the stanza in development.ini that
refers to WebError. Check your apache error log, it might tell you
what mod_wsgi is encountering as well.

http://code.google.com/p/modwsgi/wiki/DebuggingTechniques

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