TC 4.1.18, W2K Server, IIS 5

I have a question about this (copied from the TC docs):

The Host element represents a virtual host, which is an association of a
network name for a server (such as "www.mycompany.com" with the particular
server on which Catalina is running. In order to be effective, this name
must be registered in the Domain Name Service (DNS) server that manages the
Internet domain you belong to 

My question is:  Why does www.mycompany.com have to be registered in the
"DNS server" that manages the Internet domain I belong to.

In my limited knowledge, I was thinking that as long as www.mycompany.com
points to my webserver - DNS could be hosted on any DNS server.  Therefore
any request that comes to www.mycompany.com/webappname would resolve and
forward to TC.

Now, since my ISP hosts my DNS and I have TC running on the box will,
www.mycompany.com/webappname not work unless I host my own DNS?

<background>
        localhost/webappname works
        can get www.mycompany.com/webappname to work - I get a 500 error in
Explorer
        in the logs I don't see any errors - isapi.dll seems to be using my
worker from worker.properties
        - yes, I looked in the IIS logs too - can't determine what's causing
this.
</background>


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