For me the problem was arecord holding open the snd device, this was
because of the randomsound init script kicking it off to generate
entropy. try lsof | grep snd, and see anything is opening those devices
directly. Confirm, first of all with pulseaudio -vv.

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No sound in Karmic (HDA Intel ICH6 & Realtek ALC260)
https://bugs.launchpad.net/bugs/418627
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