I can verify this behavior in karmic server which has resolvconf rather than 
network-manager.
                resolvconf 1.44ubuntu1

My test was on a karmic machine.  The original install was hardy with
sequential dist-upgrades to karmic.

After the karmic upgrade was complete /etc/resolv.conf would be over-
written on boot.

The following file had my original resolv.conf file in it:
                 /etc/resolvconf/resolv.conf.d/original

/etc/network/interfaces has the following lines:
                dns-nameservers <address removed for comment>
                dns-search <domain removed for comment>

I was able to ~resolve~ my issue by adding my desired resolv.conf into the 
following:
                /etc/resolvconf/resolv.conf.d/base

My experience is that dns-nameservers in /etc/network/interfaces has
been ignored since hardy, but until karmic /etc/resolv.conf is left
untouched.

-- 
[karmic] n-m overwrites /etc/resolv.conf even when NICs are configured in 
/etc/network/interfaces
https://bugs.launchpad.net/bugs/435618
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