This is not a proper fix, but might help if the Browse... button in your
screenshot does not provide access to USC (I presume you mean Ubuntu
Software Centre).

On Lucid, Firefox recognises that a .deb file is a software package and asks if 
you want to open it with GDebi.  Try using the Browse button to navigate to 
gdebi in usr/bin
It's possible there is a bug/omission in shared-mime-info.

Not ideal, but a little better than having to save the file and then
open it with another app.

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https://bugs.launchpad.net/bugs/830911

Title:
  Firefox by default doesn't know what to do with .deb files

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