Ok, so the problem for me is the exhausting of inotify watches.
See 
https://phobosk.wordpress.com/2012/11/11/howto-fix-a-virtualbox-failed-to-access-the-usb-subsystem-error/

VBoxSVC strace out:
24511 inotify_add_watch(10, "/dev/vboxusb", 
IN_MODIFY|IN_ATTRIB|IN_CREATE|IN_DELETE) = -1 ENOSPC (No space left on device)

Fixed by increasing the limits.

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https://bugs.launchpad.net/bugs/1576768

Title:
  user can not access USB devices

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