Shouldn't they be pretty closely related, though?

It depends on from what perspective. They are related and both important in terms of the success of your website, yes, but SEO standard for "search engine optimization" and has nothing to do with usability. I website that is very well optimized might is likely to more usable than one that is not, but you can have a very, very optimized website that outperforms all of the others that is completely unusable.

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