Dear Martin,

in my opinion, it would be preferable not to deposit the coordinates of
atoms with (extremely) high B-factors (or "invisible" atoms) in PDB files.
Many users of the database, including statisticians and biologists, may not
be familiar with the concept of B-factor. When visualizing the structure
with tools like ChimeraX or PyMOL, or whenorming statistical surveys, they
might mistakenly interpret the position of these atoms as experimentally
determined.

Unfortunately, I have noticed that even some crystallographers do not fully
understand the role of B-factors, which could lead to misleading
interpretations of the data.

Best regards,

Oliviero Carugo



PS I published, years ago, something on this topic. You will find it, if
you need it.

Il giorno ven 3 lug 2026 alle ore 09:31 Alexandre Ourjoumtsev <
[email protected]> ha scritto:

> Dear all,
>
> B-factors, as well as occupancy values, are "physical" characteristics of
> a structure.
> Defining non-identified atoms with zero occupancy or with huge B-values
> (up to 10^4, as for some cryoEM models available in EMDB) has no physical
> meaning, as discussed multiple times in CCP4. Moreover (while this is not
> fully true), in overall, values of these parameters, similarly to atomic
> positions, are expected to be independent of a particular experiment.
>
> On the other hand, atoms missed in a given model are a feature of a
> particular map and not of the structure, this depends on how well this part
> of the model can be distinguished, recognized in this map (let's put aside
> not-realistic situations when one simply did not build a model within a
> clear density).
>
> Recently, we have proposed to complete a model description by one more
> parameter, a "local resolution", which is not a characteristic of the
> "physically existing, universal structure" but of the "map from which the
> deposited model was obtained". At my knowledge, this parameter has been
> formally accepted by Phenix and can be used right now.
>     This parameter allows one:
> 1) to reproduce an (experimental) variable-resolution map from an atomic
> model
> 2) for a given atom, to characterize the confidence of its parameters
> (coordinates, occupancy and ADP) obtained from a particular map
>
> Some large value (100 A?) of this parameter seems to be a better
> description of the situation that Martin reminds, and which, unfortunately,
> is frequent enough. Unless, when one, in purpose, wishes to characterize a
> highly mobile residues by large B.
>
> I understand that this would change the traditions (actually, not well
> established, as Mark and Robert confirm) but it seems to be more
> appropriate using this parameter and neither huge ADP nor zero occupancy to
> characterize the model parts poorly distinguished (totally missed) in the
> map.
>
> With best regards,
>
> Sacha Urzhumtsev
>
>
> ----- Le 2 Juil 26, à 17:13, Mark J. van Raaij <
> [email protected]> a écrit :
>
> have a look in the ccp4bb archives, this has been discussed multiple times
> without a clear conclusion
> my approach would be to keep them and let the B-factors refine to high
> values, the reason is that you know the side-chains are there and with full
> occupancy
> (the validation statistics may be worse but that is normal for low-res
> structures)
>
>
> Mark van Raaij
> Dpto de Estructura de Macromoleculas, lab B5B
> Centro Nacional de Biotecnologia - CSIC
> calle Darwin 3
> E-28049 Madrid, Spain
> tel. +34 91 585 4616 (internal 432092)
>
>
>
> On 2 Jul 2026, at 16:47, Martin Hu <
> [email protected]> wrote:
>
> Dear all,
>
> I would like to ask for some advice on the best way to handle poorly
> defined side chains for deposition of a low-resolution (~3.6 Å) X-ray
> structure.
>
> For a number of residues in my structure, there is little or no side-chain
> density, so I do not feel confident modelling the full side chains. At the
> moment, I have deleted the residues that are not supported by the electron
> density, which also gives better geometry and refinement statistics.
>
> I asked the PDBe deposition team whether they had any preference between
> deleting the unsupported side-chain atoms or keeping them with zero
> occupancy. They replied that they do not have any specific requirements, as
> long as the deposited model is appropriate.
>
> I was therefore wondering how people here would normally deal with this
> situation for PDB deposition. Would you generally:
>
> * delete the unsupported side-chain atoms,
> * keep the full side chains but set the unsupported atoms to zero
> occupancy,
> * or use another approach?
>
> I’d be interested to hear what people usually do for structures around
> this resolution.
>
> Many thanks in advance.
>
> Best regards,
>
> Martin Hu
>
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