Oliviero,

The idea of people who don't know B-factors analyzing structures gives me 
goosebumps – but you are probably right. I've seen students calculating surface 
charge maps from models that had stubs (missing side chains). And I am aware 
that most of the time, analysis and molecular dynamics programs do not take 
B-factors in account. Not to mention the number of analyses that are performed 
without a previous check of the map. I think all those are symptoms of a 
suboptimal system – mostly because there are no optimal solutions for the 
downstream processes yet, or maybe I ignore them? Not an easy task, if you 
think about it, to incorporate that 4th dimension in the calculations. If 
people who do not know B-factors are trying to analyze our structures, wouldn't 
it be better if they associate to structural biologists to get the work done 
properly? Or at least start checking the B-factors in colors, which is very 
easy to do with both PyMol and ChimeraX. My opinion at least.

Best,

---------------------------------------------------
Carlos KIKUTI, PhD
Structural Motility Team
UMR144 - CNRS - Institut Curie
12 rue Lhomond - 75005 Paris, France
[email protected]<mailto:[email protected]>



De : CCP4 bulletin board <[email protected]> au nom de Italo Carugo 
Oliviero <[email protected]>
Répondre à : Italo Carugo Oliviero <[email protected]>
Date : vendredi 3 juillet 2026 à 09:54
À : "[email protected]" <[email protected]>
Objet : Re: [ccp4bb] Side-chain deletion or zero occupancy for PDB deposition


Dear Martin,

in my opinion, it would be preferable not to deposit the coordinates of atoms 
with (extremely) high B-factors (or "invisible" atoms) in PDB files. Many users 
of the database, including statisticians and biologists, may not be familiar 
with the concept of B-factor. When visualizing the structure with tools like 
ChimeraX or PyMOL, or whenorming statistical surveys, they might mistakenly 
interpret the position of these atoms as experimentally determined.

Unfortunately, I have noticed that even some crystallographers do not fully 
understand the role of B-factors, which could lead to misleading 
interpretations of the data.

Best regards,

Oliviero Carugo



PS I published, years ago, something on this topic. You will find it, if you 
need it.

Il giorno ven 3 lug 2026 alle ore 09:31 Alexandre Ourjoumtsev 
<[email protected]<mailto:[email protected]>>
 ha scritto:
Dear all,

B-factors, as well as occupancy values, are "physical" characteristics of a 
structure.
Defining non-identified atoms with zero occupancy or with huge B-values (up to 
10^4, as for some cryoEM models available in EMDB) has no physical meaning, as 
discussed multiple times in CCP4. Moreover (while this is not fully true), in 
overall, values of these parameters, similarly to atomic positions, are 
expected to be independent of a particular experiment.

On the other hand, atoms missed in a given model are a feature of a particular 
map and not of the structure, this depends on how well this part of the model 
can be distinguished, recognized in this map (let's put aside not-realistic 
situations when one simply did not build a model within a clear density).

Recently, we have proposed to complete a model description by one more 
parameter, a "local resolution", which is not a characteristic of the 
"physically existing, universal structure" but of the "map from which the 
deposited model was obtained". At my knowledge, this parameter has been 
formally accepted by Phenix and can be used right now.
    This parameter allows one:
1) to reproduce an (experimental) variable-resolution map from an atomic model
2) for a given atom, to characterize the confidence of its parameters 
(coordinates, occupancy and ADP) obtained from a particular map

Some large value (100 A?) of this parameter seems to be a better description of 
the situation that Martin reminds, and which, unfortunately, is frequent 
enough. Unless, when one, in purpose, wishes to characterize a highly mobile 
residues by large B.

I understand that this would change the traditions (actually, not well 
established, as Mark and Robert confirm) but it seems to be more appropriate 
using this parameter and neither huge ADP nor zero occupancy to characterize 
the model parts poorly distinguished (totally missed) in the map.

With best regards,

Sacha Urzhumtsev


----- Le 2 Juil 26, à 17:13, Mark J. van Raaij 
<[email protected]<mailto:[email protected]>>
 a écrit :

have a look in the ccp4bb archives, this has been discussed multiple times 
without a clear conclusion
my approach would be to keep them and let the B-factors refine to high values, 
the reason is that you know the side-chains are there and with full occupancy
(the validation statistics may be worse but that is normal for low-res 
structures)


Mark van Raaij
Dpto de Estructura de Macromoleculas, lab B5B
Centro Nacional de Biotecnologia - CSIC
calle Darwin 3
E-28049 Madrid, Spain
tel. +34 91 585 4616 (internal 432092)


On 2 Jul 2026, at 16:47, Martin Hu 
<[email protected]<mailto:[email protected]>>
 wrote:

Dear all,

I would like to ask for some advice on the best way to handle poorly defined 
side chains for deposition of a low-resolution (~3.6 Å) X-ray structure.

For a number of residues in my structure, there is little or no side-chain 
density, so I do not feel confident modelling the full side chains. At the 
moment, I have deleted the residues that are not supported by the electron 
density, which also gives better geometry and refinement statistics.

I asked the PDBe deposition team whether they had any preference between 
deleting the unsupported side-chain atoms or keeping them with zero occupancy. 
They replied that they do not have any specific requirements, as long as the 
deposited model is appropriate.

I was therefore wondering how people here would normally deal with this 
situation for PDB deposition. Would you generally:

* delete the unsupported side-chain atoms,
* keep the full side chains but set the unsupported atoms to zero occupancy,
* or use another approach?

I’d be interested to hear what people usually do for structures around this 
resolution.

Many thanks in advance.

Best regards,

Martin Hu

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