Another possibility that no one has mentioned yet is that you may have two
accounts that are almost identical in setup. One account downloads the
messages, but has a mis-configured SMTP server (or is just a corrupt account
that needs to be deleted and re-setup). When you reply to messages
downloaded by this account the replies will be sent via this munged account,
hence the error message.

The second account is setup correctly and is your default account. Since new
outgoing messages would use this account, everything goes out fine.

-Remo Del Bello 

-- 
"Advice is a dangerous gift, even from the wise to the wise, and all courses
may run ill."
- Gildor Inglorion in "The Lord of the Rings"


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